Para La Tierra and its interns and volunteers
regularly publish scientific papers.
Read them all here.

Botany

Snelling, G. A. (2012). Familiar Ferns in a Far Flung Paradise. Pteridologist, 5(5), 312-317. (PDF 972Kb)

Invertebrates

Smith, P., Galeano, S., Ortega, C. (2025) A new country record and fifteen new departmental records for Paraguayan sphingidae (insecta: lepidoptera). Entomologia Mexicana nota cientifica 12:70-83(2025). (PDF 1.3Mb)

Borges, A., Caballero, C, de Arias, A.R., Smith, P., Owen, M., Nishi, A., Maluff, Y., Wieczorks,T. (2025). First report of the medically significant Argentine scorpion Tityus carrilloi (Buthidae) in Paraguay: Epidemiological implications amid rising regional scorpionism. Toxicon 264 108456 PDF available from Paul Smith: faunaparaguay@gmail.com

Mussini, G., Stepan, N.D., Vargas, G. (2024) Two new species of Hyalella (Amphipoda, Dogielinotidae) from the Humid Chaco ecoregion of Paraguay. ZooKeys 1191: 105–127 (2024) (PDF 11.5Mb)

Perger, R., Pett, B.L. (2022) Mazax akephaloi sp. nov.—a new Neotropical spider species resembling ‘headless’ Ectatomma ants (Araneae: Corinnidae: Castianeirinae). Zootaxa 5150(4): p579–590 (PDF 5.9Mb)

Pett, B.L. (2023) A new species of Castianeira (Araneae: Corinnidae: Castianeirinae) from central South America. Arachnology 19(6), p936–939. (PDF 3.3Mb)

Sherwood, D., Pett, B.L. (2022) Mouse spiders in the humid Chaco: two new species of Actinopus Perty, 1833 from Paraguay (Araneae: Actinopodidae). Revista Ibérica de Aracnología, 41: p75–81. (PDF 10.5Mb)

Smith, P., Kitching, I.J., Rios, S., Haxaire, J. (2022) An annotated catalogue of the Paraguayan Sphingidae (Lepidoptera). J. Insect Biodiversity 031 (2) 2022 (PDF 17.6Mb)

Pett, B.L., Wyer, R., Smith, P. (2022) Contributions to the knowledge of tiger beetles in Paraguay, with new country and departmental records (Coleoptera: Cicindelidae) Fragmenta entomologica, 54 (1): 163–170 2022 (PDF 890Kb)

Pett, B.L., Rubio, G.D., Perger, R. (2022) Grismadox gen. nov., a new Neotropical genus of ant-resembling spiders (Araneae, Corinnidae, Castianeirinae), including the description of two new species from Bolivia and Paraguay. Zoosyst. Evol. 98(1), 1–11 (PDF 5.5Mb)

Bascoules, S., Smith, P,(2021) Mutualism between frogs (Chiasmocleis albopunctata, Microhylidae) and spiders (Eupalaestrus campestratus, Theraphosidae): a new example from Paraguay. Alytes, 2021, 38: 58–63. (PDF 400Kb)

Pett, B. L. (2021). Description of a new species of Myrmecotypus (Araneae: Corinnidae: Castianeirinae), the first of its genus known from Paraguay, with an updated key to species. Zootaxa 4999(1): 77-86. PDF available from Brogan Pett: brogan.pett@outlook.com

Pett, B. L., Rubio, G. D., Stolar, C. E. (2021). A first baseline for the salticid (Araneae: Salticidae) fauna of Paraguay, with thirty-two new records and description of a new species from Paraguay and Argentina. Arachnology 18(8): 922-935.PDF available from Brogan Pett: brogan.pett@outlook.com

Pett, B.L. (2021). First description of the female of Otiothops inflatus Platnick, 1975 (Araneae: Palpimanidae). Revista Ibérica de Aracnología, 38: 103–106. (PDF 7.9Mb)

Smith, P., Lesterhuis Manomet, A. J., Rodríguez, O. (2020). The social spider Parawixia bistriata (Rengger, 1836) (Araneidae) as a potentially significant source of avian mortality in the Paraguayan Chaco, Arachnology 18 (5), 436–439 (PDF 1.94 Mb)

Dickens, J., Schönberger, D., VanCompernolle, M. (2020). Guide to the Odonata of central Ñeembucú, Paraguay: indicator species of wetland habitats. International Journal of Odonatology. 1-51. 10.1080/13887890.2020.1768157. PDF available from Jeremy Dickens: jerdickens@gmail.com

Pett, B.L., Wyer, R. (2020) First confirmed record of the genus Cybaeodamus (Araneae: Zodariidae) in Paraguay, with notes on its distribution. Revista Ibérica de Aracnología, 36: 161–162. (PDF 2.3Mb)

Pett, B.L. (2019) Eburiini (Coleoptera: Cerambycidae: Cerambycinae) in Paraguay: description of a new species, with new country and geographic records. Revista Chilena de Entomología (2019) 45 (4): 673-680. (PDF 2.3Mb)

Pett, B.L. (2019) First confirmed locality record of Gypogyna forceps Simon, 1900 (Araneae: Salticidae: Salticinae: Scopocirini) from Paraguay. Peckhamia 193.1 (PDF 726Kb)

Smith, P. (2019) Myelobia smerintha (Hubner, 1821) (insecta: lepidoptera: crambidae) also occurs in Paraguay. Boletin del Musoe Nacional de Historia Natural del Paraguay 23(1) 18-20 (PDF 2Mb)

Rios, S. D., Smith, P. (2018) Catacantha ferruginea (Draudt, 1929) [Insecta, Lepidoptera, Saturniidae], una nueva adición a la entomofauna paraguaya. Paraquaria Natural 6(2) 29-30 (PDF 2.1Mb)

Smith, P., McMahon, L. (2018) A phenodeviant specimen of the rare moth Catharisa cerina Jordan (Lepidoptera Saturniidae Hemileucinae) with asymmetrical wing venation. Revista Chilena de Entomología 44(2) 211-214 (PDF 612Kb)

Smith, P., Ríos, S. D., Petko, O. N., Smith, R. L., Atkinson, K. (2017). The hawkmoths (Insecta, Lepidoptera, Sphingidae) of Rancho Laguna Blanca, Departamento San Pedro, Paraguay with documentation of five new country records. Revista del Museo da la Plata 2(2) 77-96 (PDF 4.46Mb)

Smith, P., Rios, S. D., Petko, O. N., Kitching, I. J. (2016). Oxyteninae Jordan, 1924: first confirmed Paraguayan records (Lepidoptera: Saturniidae). SHILAP Revtisa de lepidopterologia 44 (176) 553-556 (PDF 254Kb)

Petko, O., Smith, P., Stange, L., Rios, S. D. (2016). New antlion records (Neuroptera: Myrmeleontidae) from Paraguay Insecta Mundi 0487 1-8 (PDF 338Kb)

Rios, S. D., Smith, P., Quintana, E. R., Chialchia, A. O. C., Roque, J. R. C., (2016). Revision preliminar de los saturniidae (Insecta: Lepidoptera) del departamento de Neembucu, Republica del Paraguay Boletin del Musoe Nacional de Historia Natural del Paraguay 20(1) 57-64 (PDF 2.9Mb)

Smith, P., Rios Díaz S., Petko, O., Atkinson, K. (2015). The giant butterfly moths (Lepidoptera castniidae) of Reserva Natural Laguna Blanca, Departamento San Pedro, Paraguay, with a new country record. Boletín de la Sociedad Entomológica Aragonesa 57 304–306. (PDF 3.95Mb)

Rios Díaz, S., Smith, P,, Penco F. C., González, J. M. (2015). Registros recientes de Ceretes marcelserres (Godart, [1824]) (Lepidoptera: Castniidae). Boletín del Museo Nacional de Historia Natural del Paraguay 19 29-34. (PDF 2.17Mb)

Ríos Díaz, S. D, Smith, P. (2013). Cuatro nuevas especies de Saturniidae (Insecta: Lepidoptera) para el Paraguay. Boletín del Museo Nacional de Historia Natural del Paraguay. 17 86-90. (PDF 432Kb)

Smith, P., Rios, S. D., Atkinson, K., Meikle, O., Casagrande, M. (2013). New Paraguayan Specimens and First Confirmed Phenological Data for Catharisa cerina (Jordan, 1911) (Lepidoptera: Saturniidae). Brazilian Journal of Biosciences, 11(3), 349-351. (PDF 505Kb)

Smith, P., Rios, S. D., Atkinson, K., Meikle, O. H. H., Casagrande, M. M. (2013). Almeidaia aidea Meikle and Casagrande (Lepidoptera: Saturniidae): First Record for Paraguay. Entomological News, 123(1), 38-40. (PDF 145Kb)

Gonzalez, J. M., Rios, S. D., Smith, P. (2013). Confirmation of the Presence of Geyeria uruguayana (Burmeister, 1880) in Paraguay, with notes (Lepidoptera: Castniidae). SHILAP Revta. lepid., 41(161), 105-111. (PDF 666Kb)

Rios, S. D., Smith, P. (2012). First Records of Hamadryas laodamia cramer(Lepidoptera: Nymphalidae) in Paraguay. Reportes Cientificos de la FACEN,3(1), 71-73. (PDF 544Kb)

Smith, P. (2009). Finding Cath. Southern Lepidopterists' News, 31(1), 23. (PDF 233Kb)

Herpetology

Bailliez, J., Wagner, J., Smith, P., Smith, R.L. (2024) Note on predation of Amphisbaena prunicolor in Paraguay. Herpetological Review 55(1) (JPEG 938Kb)

Smith, P., Macie, J., Hurtado, M.R., Galeano, S.,Stepan, N., Brouard, J-P., Ríos, S.D., Riveros, M.F., Smith, R.L., Owen, M., Nicolay, H., Derna, R., Burró, A.V., Lahaye, T., Cacciali, P. (2024) Twenty-eight new and significant departmental reptile records for Paraguay. Acta Zoológica Lilloana 68 (2): 347-385 (PDF 9Mb)

Smith, P., Nicolay, H., Burró, A.V., Hurtado, M.R., Smith, R., Ayala, J., Ramírez, O., Derna, R., Vera, A., Rojas, F., Saldívar, M., Ottiger-Wittig, F., Bielke, C., Ramírez, Y.C., Owen, M. (2024) Significant new reptile records for Paraguay from Boquerón, Caazapá, Canindeyú, Guairá, Itapúa, Misiones, Ñeembucú and Presidente Hayes departments. Historia Natural, Tercera Serie V14(2) 53-73 (PDF 9.3Mb)

Hiscock, M., VanCampernolle, M., Dickens, .K. (2023) Dietary competition between Hemidactylus Mabouia (Sauria: Gekkonidae) and a native anuran in Pilar, Paraguay. Herpetological Conservation and Biology 18(1):9–19. (PDF 839Kb)

Smith, p., Rojas, F. (2023) Necography by Erythrolamprus poecilogyrus (Wied, 1824) (reptilia: dipsadidae) in Paraguay. Revista Latino America de herpetologia Vol.06 No.03 (PDF 1.1Mb)

Sarvary, J., Pett, B., Atkinson, K., Smith, P. (2021) Pseudis Paradoxa Platensis (La Plata paradox frog) larval diet. Herpetological Review 52(4) p.835 (PDF 501Kb)

Hicks, G., Burro, A.V., Wang, J., Dickens, J., David, H-P., Brouard, J-P., Rios, S.D., Santacruz, J.A., Smith, P. (2022) New and significant distributionsl records of Paraguayan reptiles and amphibians from Alto Paraguay, Missiones,Ñeembucu, Presidente Hayes and San Pedro departments. Historia Natural 12(3) 21-36 (PDF 5.9Mb)

Mackenzie, C., Vladimirova, V. (2022) Food in the city: the urbanised diets of Rhinella diptycha (Anura: Bufonidae), Hemidactylus mabouia (Squamata: Gekkonidae), and Tropidurus torquatus (Squamata: Tropiduridae) in Pilar, Paraguay. Herpetology Notes 15: 523-532 (PDF 1.75Mb)

Smith, P., Ayala, J., Hicks, G., Brouard, JP. (2022) The distribution of Amphisbaena prunicolor (Cope, 1885) in Paraguay and its diagnosis from A. darwinii Duméril & Bibron, 1839.Studies on Neotropical Fauna and Environment, DOI: 10.1080/01650521.2022.2041322 (PDF 10.2Mb)

Smith, P. (2022) Rancho Laguna Blanca in Paraguay is special. Litteratura Serpentium Vol 2.1 2022 (PDF 3.1Mb)

Smith, P., Brouard, JP., Cacciali, P. (2022) A new species of Phalotris (Serpentes, Colubridae, Elapomorphini) from Paraguay. Zoosyst. Evol. 98 (1) 2022, 77–85 (PDF 4.8Mb)

Smith, P., Hicks, G., Brouard, J-P. (2021): Trilepida koppesi (Amaral, 1955) (Leptotyphlopidae) First Paraguayan record of the genus and species, with an illustrated key to Paraguayan blind-snakes (Scolecophidia). Acta Biol. Par., Curitiba, 50 (1–4): 163–174. (PDF 1.7Mb)

Smith, P., Faust, T. (2020) Possible axanthism in Rhinella ornata from Paraguay. Bol. Asoc. Herpetol. Esp. (2020) 31(2) (PDF 560Kb)

Mackenzie, C.M., Vladimirova, V. (2021) Preliminary study and first evidence of presence of microplastics in terrestrial herpetofauna from Southwestern Paraguay. Studies on Neotropical Fauna and Environment, DOI: 10.1080/01650521.2021.1895466 PDF available from Cullen Mackenzie cullenmackenzie4@gmail.com

Cacciali, P., Mee, G., von Plettenberg Laing, A., Krause, D., McLaughlin, C., Montgomery R., Smith, P. (2020). Morphological Re-Examination of the Endemic Paraguayan Snake Phalotris nigrilatus Ferrarezzi, 1993 (Serpentes: Colubridae: Elapomorphini), with Notes on Its Ecology and Conservation Status, Current Herpetology 39 (1), 28-37. (For Full PDF, email piercantropus@gmail.com)

Ríos, S.D., Bascoulès, S. Vera,A., Smith,P. (2020). Primeros ejemplares con localidades de Philodryas agassizii (Jan, 1863) [Serpentes: Dipsadidae] en la región oriental del Paraguay, Nótulas Faunísticas – Segunda Serie, 298, 1-4 (PDF 3.19 Mb)

Davis, H-P., VanCompernole, M., Dickens, J. (2020) Effectiveness and reliability of photographic identification methods for identifying individuals of a cryptically patterned toad. Herpetological Conservation and Biology 15(1):204–211. (PDF 1.4Mb)

de Franca. D.P.F., Oliveira, I.F., Gennari, D., Rocha, B., Smith, P., Scrocchi, G.J., Machado-Filho, R.R. (2020) Body-bending behaviour in snakes: new records of a poorly documented defensive behaviour. HB Vol. 9 No.1 - Notas de Historia & Distribuicao Geografica. (PDF 9.4Mb)

Smith, R. L., Smith, P. (2019). Additional observations on the nest and eggs of Stenocercus caducus (Squamata: Tropiduridae) with a report of nest usurpation, Phyllomedusa 18(2), 259-263 (PDF 1.55 Mb)

Pett, B. L., Sarvary, J., David, H. P., Smith, P., Atkinson, K. (2019) Records of morphological abnormalities of anuran limbs from Paraguay. Caudernos de Herpetologia 33(2): 79-82 (2019) (PDF 393Kb)

Smith, P. (2019) Head Triangulation in Dipsas turgida (Cope, 1868) (Dipsadidae, Dipsadinae, Dipsadini): A Potential Deimatic Behavior? Bulletin of the Chicago Herpetological Society 54(7): 142-144, 2019 (PDF 5.1Mb)

Cabral, H., Carosini, A., Smith, P. (2019) How Paraguayan snakes avoid predation and live to tell the tale. Herpetology Notes, volume 12: 513-518 (PDF 4.6Mb)

Atkinson, K., Smith, P., Dickens, J., & Lee-Zuck, C. (2018). Rediscovery of the ‘Lost’ Snake Phalotris multipunctatus (Serpentes: Dipsadidae) in Paraguay with Behavioural Notes and Reference to the Importance of “Reserva Natural” Laguna Blanca for its Conservation. Current Herpetology, 37(1), 75-80. (For Full PDF email karina@paralatierra.org)

Atkinson, K., Smith, P.. Sarvary, J. (2017). New and noteworthy snake species records (Colubridae and Dipsadidae) for the Reserva Natural Laguna Blanca, eastern Paraguay. Check List 13(1): 2027 (PDF 2.36Mb)

Cacciali, P., Scott, N. J., Ortiz, A. L. A., Fitzgerald, L. A., Smith, P. (2016). The Reptiles of Paraguay: Lioterature, Distribution, and an Annotated Taxonomic Checklist. Museum of Southwestern Biology 2016 (PDF 15.7Mb)

Smith, P., Atkinson, K., Brouard, J. P., Pheasey, H. (2016). Reserva Natural Laguna Blanca, Departmento San Pedro: Paraguay's first important area for the conservation of amphibians and reptiles? Russian Journal of Herpetology 23(1) 25–34 (PDF 111Kb)

Brouard, J. P., Manders, R., Smith, P. (2015). Elachistocleis matogrosso Caramaschi 2010, (Amphibia: Anura: Microhylidae) first records for Paraguay. Cuaderne de Herpetologia 29 (1) 97-98 (PDF 421Kb)

Smith, P., Clay, R. (2015). Six new or important departmental herpetological records for Paraguay. Paraquaria Natural 3(1) 20-22 (PDF 1.88Mb)

Brouard, J. P., Smith, P. (2014). Elachistocleis bicolor (Amphibia: Anura: Microhylidae) and Eupemphix nattereri (Amphibia: Anura: Leiuperidae): Morphological abnormalities from Reserva Natural Laguna Blanca, Paraguay. Herpetologia Brasileira 3: 85-87. (PDF 1.87Mb)

Pheasey, H., Smith, P., Brouard, J. P., Atkinson, K. (2014). Vanzosaura rubricauda(Red-tailed Vanzosaur) bifurcation and trifurcation. Herpetological Review 45: 138-139. (PDF 119Kb)

Smith, P. (2014). Physalaemus centralis (Central Weeping Frog) Vocalisation and Conservation. Herpetological Bulletin 126: 32-33. (PDF 365Kb)

Atkinson, K., Smith, R., Smith, P. (2014). Epicrates Crassus (Eastern Rainbow Boa). Herpetological Review 45(4) (PDF 185Kb)

Smith, P. (2014) The Threatened, Exploding Frogs of the Paraguayan Dry Chaco. Froglog 22(3) 111 26-28 (PDF 434Kb)

Smith, P., Cacciali, P., Scott, N., Del Castillo, H., Pheasey, H., Atkinson, K. (2014). First record of the globally-threatened Cerrado endemic snake Philodryas livida (Amaral, 1923) (Serpentes, Dipsadidae) from Paraguay, and the importance of the Reserva Natural Laguna Blanca to its conservation. Cuaderno de Herpetologia 28(1) 1-3 (PDF 825Kb)

Smith, P., Scott, N., Pheasey, H., Atkinson, K. (2013). Confirmation of the Presence of Philodryas nattereri (Steindachner, 1870), in Paraguay. Herpetozoa, 16(1/2), 91-94. (PDF 230Kb)

Smith, P., Scott, N., Cacciali, P., Atkinson, K. (2013). Rhachidelus brazili (Squamata: Serpentes): First Records from Paraguay and Clarification of the Correct Spelling of the Neneric Name. Salamandra, 49(1), 56-58. (PDF 1.79Mb)

Smith, P., Cacciali, P., Kallberg, A., Atkinson, K., Pheasey, H. (2013). Reptilia, Squamata, Serpentes, Lygophis paucidens (Hoge, 1952): First Records for Paraguay. Check List, 9(1), 131-132. (PDF 620Kb)

Smith, P., Cacciali, P., Atkinson, K., Pheasey, H., Motte, M. (2012). New Distributional Records of Amphibians for Departamento San Pedro, Paraguay (Amphibia). Check List, 8(5), 903-907. (PDF 2.45Mb)

Smith, P., Cacciali, P., Atkinson, K., Kallberg, A., Pheasey, H. (2011). Nuevos Registros de Gymnophthalmidae (Reptilia: Sauria) en la Reserva Natural Laguna Blanca, Departamento San Pedro, Paraguay y una Clave para las Especies Paraguayas. Notulas Faunisticas, 81, 1-6. (PDF 1.86Mb)

Ichthyology

VanCompernolle, M., Pett, B.L. (2020) New records of Riggia cryptocularis (Isopoda: Cymothoidae) parasitizing two Odontostilbe species (Characiformes: Characidae) in the Paraguay River Basin, Paraguay, with a review of Riggia host species. Pan-American Journal of Aquatic Sciences, 15(3): 204-217 (PDF 816Kb)

Dickens, K. (2019). Annotated checklist to the fish of Laguna Blanca, San Pedro, Paraguay. Aqua, International Journal of Ichthyology, 25(4), 155-178. (for full PDF email jerdickens@gmail.com)

Mammalogy

Smith, P., Rios, S.D. (2026) Nomenclatorial and geographical notes on some Ctenomys (Rodentia: Ctenomyidae) from Paraguay and northern Argentina associated with Julio Rafael Contreras. Mammalia 2026; aop (PDF 459Kb)

Smith, P., Derna, R. (2026) A new Paraguayan departmental record for Lasiurus (Dasypterus) ega (Chiroptera: Vespertilionidae) with an updated distribution map for the species and comments on migratory status. Boletin de Museo Nacional Historia Natural del Paraguay Vol. 30, Art. e2026007 (PDF 1.7Mb)

Smith, P., Galeano, S., Rojas, F. (2026) Chital, Axis axis Erxleben, 1777 (Mammalia: Cervidae), in southern Paraguay. Boletin de Museo Nacional Historia Natural del Paraguay Vol. 30, Art. e2026004 (PDF 1.6 Mb)

Smith, P., Ayala, J., Kobrinsky, W.Y., Owen, M., Smith, R.L. (2026) Significant new records of large and medium-sized mammals in southern Paraguay. Mammalia æquatorialis, 8(2): 39–44, 2026 (PDF 2.1Mb)

Smith, P., Owen, M.T. (2026) The Paraguayan micro-endemics Ctenomys paraguayensis Contreras, 2000 and Ctenomys pilarensis Contreras, 1993 (Mammalia: Ctenomyidae) are morphologically indistinguishable. Mammal Research (2026) 71:47

Taylor, R.V., Hurtado, M.R., Owen, M., Smith, R.L. (2024) First record of twins in wild black and gold howler monkeys (Alouatta caraya). Neotropical Primates 30(2) (PDF 2.2Mb)

Smith, P. (2025) A Case Report: Syndactyly in a Wild, Roadkill, Feline: Geoffroy’s Cat Leopardus geoffroyi (Mammalia: Felidae). Research in Veterinary Science and Medicine 2026 6(1) (PDF 497Kb)

Smith, P. (2025) Status, conservation and distribution of Paraguayan Hystricomorpha2, Echimyidae (rodentia). FACENA 35 (1). 2025 (PDF 989Kb)

Weksler et al. A comprehensive survey of mammal collections and genetic resources in South America: challenges and directions. Biological Journal of the Linnean Society, 2025, 146(1). For full PDF email rebecca@paralatierra.org

Ortega, C., Segovia, P., Smith, P. (2025) Some notes on Tayra Eira barbara (Linnaeus, 1758) (mustelidae: guloninae) in Paraguay. Revista Nordestina de Zoologia, 15(1): 33-43. (PDF 1.6Mb)

Smith, P. (2025) Status and distribution of Paraguayan Sciuridae (Rodentia) and Leporidae (Lagomorpha). Russian journal of theriology 24 (1): 37–48 (PDF 632Kb)

Smith, P., Smith, R.L., Owen, M.T. (2025) Some natural history notes on Paraguayan armadillos (Xenarthra, Chlamyphoridae) Tolypeutes matacus and Euphractus sexcinctus. Xenarthra 26 (2025): e2601 (PDF 1.9Mb)

Smith, P. (2025) Status and distribution of Paraguayan mammalia hystricomorpha 1. erethizontidae and chinchillidae. Zoodiversity, 59(2): 120–135 (PDF 1.7Mb)

Owen, R.D., Smith, P. (2025) Paraguayan mammals and mammalogy. Sarem Series C V1 (PDF 3.7Mb)
High resolution version (PDF 15.3Mb)

Smith, P., Barreto, M.B., Vargas, G., Owen, M., Amarilla-Stevens, H., Ruedi, M., Rojas, F., Owen, R.D. (2024) New departmental records for Paraguayan bats (Mammalia: Chiroptera). Journal of Bat Research & Conservation V17(1) 2024 (PDF 1.3Mb)

Supplementary information (PDF 1.5Mb)

Smith, P. (2024) Structural abnormalities in Paraguayan southern three-banded armadillos Tolypeutes matacus (Chlamyphoridae). Xenarthra 25: e2502 (PDF 2.5Mb)

Smith, P., Rios, S.D. (2024) Status and distribution of Paraguayan Procyonidae and Mephitidae (Mammalia: Carnivora). DOI: 10.4081/nhs.2024.727 (PDF 2.1Mb)

Smith, P., Ayala, J., Smith, R.L., Cronin, B., Barrios, F., Narvaez, H., Zickgraf, O., Resquin, J.J., Armadans, C.F., Rios, S.D. (2024) New Paraguayan records of Xenarthra with a review of the species present in Ñeembucú department. Xenarthra 24: e2401 (PDF 4.29Mb)

Smith, P., Hicks, G. (2023) Patterns of fluorescence in a collection of Paraguayan Didelphidae. Brazilian Journal of Mammalogy, 2023; v.92: e92202389 (PDF 907Kb)

Smith, P. (2024) Bertoni´s Mammals: An annotated interpretation of the 1939 Catálogos Sistemáticos de los Vertebrados del Paraguay. Zoological Bibliography 2024 7(1) (PDF 9Mb)

Smith, P., Marshall, C., Pullan, C., Rojas, F., Owen, R.D. (2023) Notes on Pearson’s Chaco Mouse Andalgalomys pearsoni (Myers, 1977) Rodentia: Cricetidae: Sigmodontinae) in the Paraguayan Chaco. Galemys 35, 2023 (PDF 755Kb)

Jordan, K.C., Hicks, G., O'Leary, F. (2023) Use of abandoned nests of golden-winged cacique Cacicus chrysopterus (Vigors, 1825) by Chacoan mouse opossum Cryptonanus chacoensis (Tate, 1931) Notas sobre Mamíferos Sudamericanos 2023, vol. 5 (PDF 2.9Mb)

Smith, P., Smith, R.L. (2023) Natural history notes on Pampas Fox Lycalopex gymnocercus (Mammalia: Carnivora: Canidae) in the Paraguayan Chaco. Sitientibus série Ciências Biológicas 23: 10.13102/scb9149 (PDF 974Kb)

Smith, P. (2023) Did Azara discover Myotis simus or Myotis midastacus (Mammalia: Chiroptera: Vespertilionidae) at least a century before their formal description? ZOOLOGIA 40: e22046 (PDF 803Kb)

Owen, R.D., Gustafson, A.W., Smith, P. (2023) Recent photographic records, a new specimen, and a review of the distribution and habitat associations of Caluromys lanatus (Mammalia, Didelphidae) in Paraguay. Notas sobre Mamíferos Sudamericanos 2023, vol.5 (PDF, 2.6Mb)

Smith, P. (2022) Status and distribution of Paraguayan Mustelidae. Actazoologica cracoviensia, 65(1-2) 2022 (PDF 1.2Mb)

Smith, P. (2022) Status and distribution of Paraguayan canids. Canid Biology & Conservation 25: 1-12 2022 (PDF 826Kb)

Smith, P., Teta, P. (2022) Confused identities: the case of the chauve–souris septième or chauve–souris brun–blanchâtre of Azara (1801). DOI: 10.12933/therya_notes-22-79 (PDF 148Kb)

van Lunteren, P., Groenewold, M.A., Pozsgai, G, Sravary, J. (2021) Does voluntary wheel running exist in Neotropical wild mammals? Acta Ethologica 24, 23-30

Smith, P. (2020). What is Lutra paranensis rengger, 1830? IUCN Otter Spec. Group Bull. 37(1) 2020 (PDF 1Mb)

Smith, R.L., Smith, P. (2019). Wild observation of infanticide and cannibalism by Azara’s Agouti ( Dasyptocta azarae ) (Mammalia: Dasyproctidae) in Paraguay, Ethology 125(11). (For Full PDF, email faunaparaguay@gmail.com)

Rios, S.D., Diaz, C.R., Smith. P. (2019) Reporte de un ejemplar albino de Tamandua tetradactyla (Linnaeus, 1758) (Pilosa, Myrmecophagidae) en el suroeste de Paraguay. Edentata 20 (2019): 35–38. (PDF 697Kb)

Smith, P. (2019) An observation of water-bathing in Chaco Peccary. Suiform Soundings 18(1) (PDF 189Kb)

Owen, R., Smith, P. (2019) Mammals of the Mbaracayú Forest Natural Reserve, Canindeyú Department, Represented in Scientific Collections. Revista de la Sociedad Científica del Paraguay June 2019 volume 24(1) (PDF 219Kb)

Smith, P., Rios, S. D. (2018) Distribution and status of Paraguayan Xenarthra. Edentata 19 1–29 (PDF 3.5Mb)

Owen, R. D., Sánchez, H., Atkinson, K., McMahon, L., Jonsson, C. B. (2018). New and noteworthy records of rodents (Mammalia, Rodentia, Cricetidae and Echimyidae) from Paraguay. Check List 14 (5) 721–730. (PDF 5Mb)

Pheasey, H., Smith, P., Atkinson, K. (2018) Observations on the diet and behaviour of captive Monodelphis (Mygalodelphys) kunsi Pine, 1975 (Didelphimorphia Didelphidae) from Paraguay. Mammalogy Notes | Notas Mastozoológicas Sociedad Colombiana de Mastozoología 4 (2) (PDF 4.6Mb)

Smith, P., Owen, R. D. (2017). Calyptophractus retusus (Cingulata: Dasypodidae). Mammalian Species 49(947): 57–62 (PDF 13Mb)

Smith, P., Owen, R. D. (2016). Marmosa constantiae (Didelphimorphia: Didelphidae). Mammalian Species 48(941): 123–129 (PDF 502Kb)

Smith, P., (2015). Un tatu bolita Tolypeutes matacus (Desmarest, 1804) con una amputación de la pierna delantera de Paraguay. Paraguay Biodiversidad 2(8): 35-38 (PDF 688Kb)

Owen, R. D., Smith, P., López-González, C., Ruíz Díaz, M. (2014). First records of two species of bats (mammalia: Chiroptera: Emballonuridae and Phyllostomidae) from Paraguay. Boletín de la Sociedad Zoológica del Uruguay 23: 67-73. (PDF 157Kb)

Smith, P., Owen, R. D., Del Castillo, H., Ortiz, M. L., Cabrera, A. (2013). Historical and recent records of Greater Grison Galictis vittata in Paraguay, with nomenclatural comments. Small Carnivore Conservation 49: 43–47, (PDF 1.2Mb)

Smith, P. (2012). Assessing the Assessment: The Relevance of the 2006 Paraguayan Mammal Red List to the Reality of Xenarthra Conservation in 2012. Edentata, 13, 18-28. (PDF 581Kb)

Smith, P., Pheasey, H., Atkinson, K., Ramakers, J., Sarvary, J. (2012). The Didelphimorphia (Didelphidae) of Reserva Natural Laguna Blanca, Departamento San Pedro, Paraguay. Acta Zoológica Lilloana, 56(1-2), 141–153. (PDF 1.39Mb)

Smith, P., Pheasey, H., Atkinson, K., Miller, J. (2012). Records of the Phyllostomine Bats Tonatia bidens (Spix, 1823) and Lophostoma silvicolum (d´Orbigny, 1836) (Chiroptera, Phyllostomidae) Associated with Human Dwellings in Paraguay. Chiroptera Neotropical, 18(2), 1139-1143. (PDF 177Kb)

Smith, P., Owen, R., Atkinson, K., del Castillo, H., Northcote-Smith, E. (2011). First Records of the Southern Naked-Tailed Armadillo Cabassous unicinctus(Cingulata: Dasypodidae) in Paraguay. Edentata, 12, 53-57. (PDF 1.89Mb)

Primatology

Blood, R.A., Smith, R.L. (2026). Preliminary observations into understanding the effects of urbanization on the black-and-gold howler monkeys (Alouatta caraya) in Pilar, Ñeembucú, Paraguay. Journal of Urban Ecology. 12. 10.1093/jue/juag005.

Smith, R.L., Lusseau, D. (2023). The Hooded Capuchin Monkey (Sapajus cay) is Vulnerable in Paraguay and at Least Near Threatened Globally According to Red List Criteria. International Journal of Primatology. 45. 10.1007/s10764-023-00400-8

Smith, R.L., Santacruz, J.D. (2024). Paraguay's First Primatology and Environmental Education Training Course for Forest Guards. Primate Conservation. 38. 1-9.

Smith, R.L., Santacruz, J.D. (2024). Primate focused conservation education in Paraguay: Doing something Para La Tierra. Folia Primatologica. 95. 1-10. 10.1163/14219980-bja10039.

Taylor, R.V., Hurtado, M.R., Owen, M., Smith, R.L. (2024). First record of twins in wild black and gold howler monkeys (Alouatta caraya). Neotropical Primates. 30. 10.62015/np.2024.v30.790.

Blood, R.A., Smith, R.L. (2026) Preliminary observations into understanding the effects of urbanization on the black-and-gold howler monkeys (Alouatta caraya) in Pilar, Neembucu, Paraguay. Journal of Urban Ecology, 2026, 12(1) (PDF 1.9Mb)

Hurtado, R.H., Owen, M.T. (2025) Observation of an apparently human-directed agonistic throat display in two adult males of Alouatta caraya (Atelidae: Primates) in Pilar, Paraguay. Mammalogy Notes ISSN 2382-3704 (PDF 287Kb)

Taylor, R.V., Hurtado, M.R., Owen, M.T., Smith, R.L. (2023) A record of diurnal birth and placentophagia in an urban-dwelling female black and gold howler monkey (Alouatta caraya, Primates: Atelidae) in the City of Pilar, south-west Paraguay. Mammalogy Notes 2023, 9(2), 398 (PDF 876Kb)

Smith, R.L., Santacruz. J.D.A., Smith, P. (2022) The challenges of primate research and conservation in Paraguay. Neotropical Primates 28(1-2), December 2022. (PDF 717Kb)

Smith, R.L., Rebergen, K., Payne, C., Megaponas, E., Lusseau, D. (2022) Dietary plasticity of an umbrella species (Sapajus cay) in a biodiversity hotspot: applying ecological traits to habitat conservation in the Upper Paraná Atlantic Forest. Folia Primatologica 0 (2022) 1-16

Alesci, M., Smith, R.L., Ayala Santacruz , J.D., Camperio Ciani, A. (2022) Attitudes towards urban black and gold howler monkeys (Alouatta caraya) in the city of Pilar, Ñeembucú, southwest Paraguay. Primates 63, 161–171. (PDF 888Kb)

Smith, R.L., Briggs, E., Smith, P. (2020) Unusual behavioural responses of the South American coati Nasua nasua (Carnivora: Procyonidae) to hooded capuchins Sapajus cay (Primates: Cebidae) in the Paraguayan Upper Paraná Atlantic Forest. Neotropical Primates 26(3): 23-29. (PDF 144Kb)

Wellian, J., Smith, R.L. (2021) Risk awareness of black-and-gold howler monkeys living in an urban environment in south-west Paraguay. Journal of Urban Ecology, 2021, 1–8 (PDF 637Kb)

Smith, R.L., Blood, R., Smith, P. (2021) Facial malformations, including cleft lip and palate, in wild primates in Paraguay. J Med Primatol. 2021;00:1–4. PDF available from Rebecca Smith rebecca@paralatierra.org

Smith, P., Rios, S.D., Smith, R.L. (2021) Paraguayan Primatology: Past, Present and Future. Primate Conservation 2021(35) (PDF 1.5Mb)

Kane, J., Smith, R.L. (2020). Bertiella sp. (Meyner, 1895) infection of Alouatta caraya (Humbolt, 1812) in urban and natural environments in Ñeembucú, southwest Paraguay (For full PDF, email rebecca@paralatierra.org)

Culot, L. et. al. (2018) A dataset of communities and occurrences of primates in the Atlantic forests of South America. Ecological Society of America website. (PDF 3Mb)

Smith, R. L., Hayes, S. E., Smith, P., Dickens, J. K. (2017). Sleeping site preferences in Sapajus cay Illiger 1815 (Primates: Cebidae) in a disturbed fragment of the Upper Parana´ Atlantic Forest, Rancho Laguna Blanca, Eastern Paraguay. Primates DOI 10.1007/s10329-017-0626-7 (PDF 470Kb)

Smith, R. L. (2017). An observation of an unusual human-directed threat display by Sapajus cay Illiger, 1815 (Cebidae: Primates) in a fragment of Upper Paraná Atlantic Forest, eastern Paraguay. Mammalogy Notes, Sociedad Colombiana de Mastozoología 4 (1) 15-16 (PDF 386Kb)

Smith, R. L., Payne, C. J. (2017). Consumption of the bark of Albizia niopoides (Mimosaceae) by Sapajus cay Illiger, 1815 (Cebidae: Primates) in the Atlantic Forest fragment of Rancho Laguna Blanca in Eastern Paraguay. Mammalogy Notes, Sociedad Colombiana de Mastozoología 4 (1) 17-20 (PDF 598Kb)

Smith, R. L. (2016). Preliminary observations of complex object manipulation in a wild population of Sapajus cay Illiger 1815 (primates: Cebidae) in a fragment of upper Paraná Atlantic Forest, Rancho Laguna Blanca, eastern Paraguay. Acta Ethologica (PDF 613Kb)

Smith, R. L., Briggs, E. S. (2016). Using camera traps to determine group demography in a Paraguayan population of Sapajus cay. Neotropical Primates 22(2) 81-88 (PDF 290Kb)

Ornithology

Smith, P., Smith, R.L. (2026) Use of immature fruits of Acrocomia (Arecaceae) palms for hydration by Red-and-green Macaw Ara chloroptera (Aves: Psittacidae) and Hooded Capuchin Sapajus cay (Mammalia: Primates: Cebidae), Biodiversity, DOI: 10.1080/14888386.2026.2701023 PDF available from Paul Smith faunaparaguay@gmail.com

Doerksen, J., Smith, P., Recalde, L., Sergio, D. (2025) On the presence of Oxyura vittata (Aves, Anserifomes, Anatidae) in Paraguay. Revista de Zoología. 39: 46-50. (PDF 469Kb)

Smith, P., Rice, N.H. (2025) The correct name of the southern subspecies of Squirrel Cuckoo Piaya cayana (Cuculidae). Avian Systematics 2025 3 (1): 1–7 (PDF 3Mb)

Doldan, L., Irala, R., Rios, S.D., Ortega, C., Lowen, D.O., Smith, P. (2025) Status and distribution of the White-throated Kingbird Tyrannus albogularis (Passeriformes, Tyrannidae) in Paraguay. Historia Natural Series 3 V15(2) 71-77 (PDF 2.3Mb)

Smith, P., Paredes, J., Stelmach, H., Plessas, J., Daniels, C., Clay, R. (2025) Some colour aberrations in Paraguayan birds resulting in areas of white plumage. Revista Ecuatoriana de Ornitología 11(1) (PDF 1.8Mb)

Smith, P., Fisher, J., Naumoff, R., Rojas, F. (2024) Observation of attempted courtship feeding in the Chaco Owl Strix chacoensis (Aves: Strigidae). Revista Catalana d’Ornitologia 40:44-46, 2024 (PDF 3.4Mb)

Smith, P., Secelle, A., Carpentier, G., Plessas, J., Rojas, F. (2024) Observations on Black-legged Seriema, Chunga burmeisteri, and five other birds from the Paraguayan Dry Chaco. Boletin del Museo Nacional de Historia Natural del Paraguay V28, 16-21 (PDF 3.2Mb)

Smith, R.L., Smith, P. (2024) Observation on breeding of Silky-tailed Nightjar Antrostomus sericocaudatus sericocaudatus (Caprimulgidae) from Paraguay. Acta Biológica Catarinense 11(1):42-45 (PDF 532Kb)

Smith, P. (2023) What is Rallus rytirhynchos Vieillot, 1819? The true identity of Azara’s (1805) No. 372 Ypacahá del Pardo.Iheringia, Série Zoologia, 113: e2023016 (PDF 305Kb)

Smith, P. (2023) The overlooked nomenclatural history of the Fernando Po Mannikin Spermestes bicolor poensis (Fraser, 1843) (Estrildidae). bboc.v143i3.2023.a3 (PDF 425Kb)

Dickinson, E.C., Smith, P., Sieradzki, A., van der Mije, S., Kamminga, P. (2022) The history of the name Strix pumila and its authorship. Bull. B.O.C. 2022 142(3) (PDF 1.7Mb)

Hicks, G., Dickens, J., Torres, C., Walker, E., Rios, S.D., Doldan, L., Torre, I., Lesterhuis, A.J., del Castillo, H., Rodriguez, O., Smith, P. (2022) The Avifauna of the Paraguayan department of Neembucu. Boletín del Museo Nacional de Historia Natural del Paraguay 26/A: 1-54 (PDF 12.1Mb)

Smith, P., Sidelink, M., Amez, I.M., Rios, S.D. (2021) Some ornithological observations of interest from Área de Reserva para Parque Nacional San Rafael, Itapúa department, Paraguay. Paraquaria Nat. 8: 15-18 (PDF 1.9Mb)

Vallotton, L., Smith, P. (2021) Paraguayan bird specimens in the Museum of Natural History, Geneva, Switzerland. Boletín del Museo Nacional de Historia Natural del Paraguay 25/2: 1-92 (PDF 2.5Mb)

Smith, P., Centron, S. (2021) Nesting Observations on a South American Subspecies of Least Grebe (Tachybaptus dominicus brachyrhynchus) Waterbirds, 43(3-4): 348-352 (PDF 428Kb)

Smith, P. (2021) Black-bellied Seedeater Sporophila melanogaster (Aves: Thraupidae): the first Paraguayan record. Boletín del Museo Nacional de Historia Natural del Paraguay 25/2: 106-108 (PDF 856Kb)

Smith, P., Atkinson, K., Davies, Y.E. (2021) Paraguyan bird species in the Ibis collection. Boletín del Museo Nacional de Historia Natural del Paraguay 25: 1-51 (PDF 2Mb)

Paredes, J., Clay, R.P., Clay, M., Smith, P. (2021) Broad-winged Hawk Buteo platypterus (Aves: Accipitridae): An addition to the Paraguayan Avifauna. Paraquaria Natural 8: 1-3 (PDF 984Kb)

Smith, P. (2021) Notes on vultures (Cathartidae) in Paraguay with a supporting bibliography. Vulture News Vol. 79 (PDF 763Kb)

Smith, P. (2021) What is Azara’s (1805) No. 193 Suiriri pardo amarillo menor? Holotipus rivista di zoologia sistematica e tassonomia II (1) 2021: 3-10 (PDF 1.1Mb)

Dickens, J.K., Bitton, P., Bravo, G.A., Silveira, L.f. (2021) Species limits, patterns of secondary contact and a new species in the Trogon rufus complex (Aves: Trogonidae), Zoological Journal of the Linnean Society, 2021;, zlaa169

Smith, P., Clay, R.P. (2021) The identity of Félix de Azara’s “Alondras” and implications for Neotropical pipit nomenclature (Aves, Motacillidae: Anthus). Zootaxa 4942 (1) PDF available from Paul Smith faunaparaguay@gmail.com

Smith, P., del Castillo, H. (2021) Aramides saracura and Aramides cajaneus do not have a parapatric distribution, at least not in Paraguay: A reply to Marcondes and Silveira (Zookeys 500, 2015). Neotropical Biodiversity 7(1): 57060 (PDF 1.3Mb)

Smith, P. (2020). Azara's No. 12 Gavilán de estero chorreado and Circus albicollis Vieillot, 1816 do not refer to the Long-winged Harrier Circus buffoni (Gmelin, 1788). North-Western Journal of Zoology 16(2): 225-228 (PDF 151 Kb)

Guest, K., Sarvary, J., Smith, P. (2020). New and noteworthy bird records from the Cerrado de Laguna Blanca IBA, San Pedro department, Paraguay. Boletín del Museo Nacional de Historia Natural del Paraguay 24: 75-83 (PDF 4.22 Mb)

Smith, P. (2020). The identity of Azara´s (1805) No. 246 Trepador remos y cola roxos (Aves: Furnariidae), Zootaxa 4766(4), 594-598. (For full PDF email faunaparaguay@gmail.com)

Smith, P. (2020) Azara’s spinetails II: what is No. 239 “Cola aguda cola sanguina”? Ornithology Research 28: 191–194 (PDF 628Kb)

Smith, P. (2020). Azara’s No. 21 “Gavilán pardo obscuro” is not a Short-tailed Hawk Buteo brachyurus. Neotropical Biology and Conservation 15(4): 409–414. (PDF 139Kb)

Lesterhuis, A.J., Smith, P., Clay, R.P. (2020) First record of Himantopus mexicanus mexicanus and notes on breeding of H. m. melanurus in Paraguay. Wader Study 127(2): 162–164. (PDF 1.2Mb)

Smith, P. (2020) Azara's no. 243 ‘Trepador de pico corto’ is a Lesser Woodcreeper Xiphorhynchus fuscus. BBOC v140i1.2020.a5 (PDF 2.1Mb)

Smith, P., Rios, S.D., Doldán, L., Ruiz, R. (2019). Bill deformation in five species of Paraguayan birds, Revista Chilena de Ornitología 25(1): 3-8 (PDF 1.73 Mb)

Rodriguez, O., Castillo, L., Smith, P., Del Castillo, H. (2019). Status and distribution of Paraguayan macaws (Aves: Psittacidae) with a new country record, Papéis Avulsos de Zoologia (São Paulo), 59(60), 1-10 (PDF 3.24 Mb)

Smith, P., Easley, K. (2019) Status and distribution of Paraguayan “Black Tyrants” (Tyrannidae: Knipolegus) with a remarkable new country record. Avocetta 43: 149-158 (PDF 1.6Mb)

Smith, P., Johnson, K.P. (2019) Dew-bathing in the Atlantic Forest endemic Robust Woodpecker Campephilus robustus (Lichtenstein, 1823) (Aves: Picidae). Turkish Journal of Zoology 43: 400-402 (PDF 1.1Mb)

Fast, M., Rios, S.D., Clay, R.P., Smith, P. (2019) Sporophila nigricollis (aves, thraupidae) in Paraguay: photographic documentation and comments on its status in the country. Boletin del Museo Nacional de Historia National del Paraguay 23(1) 14-17 (PDF 945Kb)

Smith, P., Ríos, S.D. (2019) Re-thinking wing-flashing in Mimidae, with new reports from Paraguay, Ethology Ecology & Evolution. (For full PDF email faunaparaguay@gmail.com)

Smith, P., Morris, P. (2018) Crypsis in two species of tinamou (Tinamiformes: Tinamidae): roosting Solitary Tinamous [Tinamus solitarius (Vieillot, 1819)] probably mimic owls. Boletin do Museo Paraense Emílio Goeldi. Ciencias Naturais 13 (3) 467-470 (PDF 418Kb)

Smith, P. (2018) The identity of two of Azara’s “mystery” waterbirds. Caldasia 40 (2) 383-389 (PDF 300Kb)

Mello, L. J. R., Rodriguez, O., Smith, P. (2018) Confirmation that Little Cuckoo Coccycua minuta occurs in Paraguay. Revista Venezolana de Ornitología 8 59–60 (PDF 1.8Mb)

Argentini, N., Smith, P., Rodriguez, O., del Castillo, H., Rios, S. D. (2018) The Azure Jay Cyanocorax caeruleus (Aves: Corvidae) in Paraguay, with restriction of the type locality. Revista Brasileira de Ornitologia 26(3): 210–213 (PDF 1.45Mb)

Smith, P. (2018) Azara’s spinetails (Aves: Furnariidae). The identity of No. 236 Chiclí and No. 237 Cógogo. Bonn Zoological Bulletin 67 (2): 171–174 (PDF 295Kb)

Smith, P. (2018) The identity of the mysterious ‘Azara’s Parakeet’ Sittace flaviventris Wagler, 1832. Anais da Academia Brasileira de Ciências 90(3): 2851-2854 (PDF 835Kb)

Lesterhuis, A. J., Clay, R. P., Smith, P. (2018) New and noteworthy records of waterbirds from Paraguay. Revista Brasileira de Ornitologia 26(1): 65–73 (PDF 1.1Mb)

Smith, P., Cacciali, P., Carmagnola, C. (2017) A Paraguayan yellow-hooded blackbird (Chrysomus icterocephalus) and a Paraguayan blackbird with a yellow hood! Boletin del Museo Nacional de Historia National del Paraguay 21 (2) 83-86 (PDF 2Mb)

Smith, P. (2017). The identity of Azara's No. 315 “Ibiyau de cola extraña” and No. 316 “Ibiyau anónimo” (Aves: Caprimulgidae) Zootaxa 4337 (4): 595–599 (PDF 171Kb)

Smith, P, Rios, SD. (2017). Novel plumage aberrations in Paraguayan non-Passerine Birds, and the definition of a new plumage aberration unique to Psittacidae. Biodiversity and Natural History 3 (1) 24-28 (PDF 1.09Mb)

Smith, P. (2017). Distraction display of the southernmost subspecies of the Golden-crowned Warbler Basileuterus culicivorus azarae (Aves: Parulidae) and a need for more detailed documentation of paratreptic behaviours. Acta Ethologia 20:187–189 (PDF 263Kb)

Bodrati, A., Smith, P., del Castillo, H., Trejo, A. (2017). The Owls of Paraguay. Neotropical Owls, DOI 10.1007/978-3-319-57108-9_18 (PDF 693Kb)